Ayed, Fakher, 1National Agronomic Institute of Tunisia, 1082 Tunis, University of Carthage, Tunisia 2Technical Centre of Organic Agriculture, 4042 Chott-Meriam, Sousse, Tunisia 3UR13AGR09-Integrated Horticultural Production in the Tunisian Centre-East, Regional Research Centre on Horticulture and Organic Agriculture, University of Sousse, 4042, Chott-Meriem, Tunisia, Tunisia